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This is one reason why I have PS7 alongside CS in my computer. I find Image Ready tamer to handle.

I tried MrTom's remedy in PS IR 7 awhile back when he posted it and worked. But trying an hour ago in CS gave me the result Sam got.

The masks shouldn't have any effect with the entire ani. If a layer in the frame is off, it won't show in succeding frames in the sequence.  And I find nothing wrong with the masks in each layer - they're correctly positioned. Except that on closer inspection, the masks needed a bit of retouch to eliminate those white artifacts.  I guess it's the result of using the quick selection tool rather than the pen tool.

 @IamSam.....  I've still got work to do with another apps and can't afford to shut it down as it's a big file work.

Can you try what I suggested earlier - deleting the animation timeline, save the file, close then reopen and reconstruct the animation with either Do Not Dispose or Automatic on?  Deleting the frame timeline and saving should result in a much smaller psd file size. Let's see what happens.....

Thanks.

edit.....   oooops..... Sam was here but now is outta the building..... .